Hi cousin! Not sure i can be of any help because i found the line above William Grancier Mullins to be murky, with conflicting information (including from the website you mentioned. My line goes, like yours, from William G. thru John H. To David D, then my line goes thru daughter Neurany Mullins.
I did take the ancestry.com dna test and have matches with other members to William Grancier Mullins, but haven't gained any more useful information on William's parents.
I've read lots of stuff pointing to Abraham Des Moulins, but i haven't seen any hard documentation on that (other than people repeating it) let alone which woman was his wife/mother of William G!
Hope others with more information/documentation answer your query because it is a question i have also!